yat-gandha-mātrāt - simply by the scent of that elephant; harayaḥ - lions; gaja-indrāḥ - other elephants; vyāghra-ādayaḥ - ferocious animals like tigers; vyāla-mṛgāḥ - other ferocious animals; sakhaḍgāḥ - rhinoceroses; mahā-uragāḥ - big, big serpents; ca - also; api - indeed; bhayāt - because of fear; dravanti - running away; sa - with; gaura-kṛṣṇāḥ - some of them white, some of them black; sarabhāḥ - sarabhas; camaryaḥ - also camarīs.

Simply by catching scent of that elephant, all the other elephants, the tigers and the other ferocious animals, such as lions, rhinoceroses, great serpents and black and white sarabhas, fled in fear. The camarī deer also fled.