madhura-rase - in the mellow of conjugal love; bhakta-mukhya - the chief devotees; vraje - in Vṛndāvana; gopī-gaṇa - the gopīs; mahiṣī-gaṇa - the queens in Dvārakā; lakṣmī-gaṇa - the goddesses of fortune in Vaikuṇṭha; asaṅkhya gaṇana - of innumerable reckoning.

“The chief devotees in conjugal love are the gopīs in Vṛndāvana, the queens in Dvārakā and the goddesses of fortune in Vaikuṇṭha. These devotees are innumerable.