ākāra - of features; svabhāva - of natures; bhede - with differences; vraja-devī-gaṇa - the gopīs; kāya - of Her body; vyūha - of expansions; rūpa - having the form; tāṅra - of Her; rasera - of mellows; kāraṇa - instruments.

The Vraja-devīs have diverse bodily features. They are Her expansions and are the instruments for expanding rasa.