srotram -- the sense of hearing; tvak -- the sense of touch, experienced upon the skin; darsanam -- sight; ghranah -- smell; jihva -- the sense of taste, experienced upon the tongue; iti -- thus; jnana-saktayah -- the knowledge-acquiring senses; vak -- speech; pani -- the hands; upastha -- the genitals; payu -- the anus; anghrih -- and the legs; karmani -- the working senses; anga -- My dear Uddhava; ubhayam -- belonging to both these categories; manah -- the mind.

Hearing, touch, sight, smell and taste are the five knowledge acquiring senses, My dear Uddhava, and speech, the hands, the genitals, the anus and the legs constitute the five working senses. The mind belongs to both these categories.