tayā - by the doe; pepīyamāne - being drunk with great satisfaction; udake - the water; tāvat eva - exactly at that time; avidūreṇa - very near; nadataḥ - roaring; mṛga-pateḥ - of one lion; unnādaḥ - the tumultuous sound; loka-bhayam-kara - very fearful to all living entities; udapatat - arose.

While the doe was drinking with great satisfaction, a lion, which was very close, roared very loudly. This was frightful to every living entity, and it was heard by the doe.