śrotram - the sense of hearing; tvak - the sense of touch, experienced upon the skin; darśanam - sight; ghrāṇaḥ - smell; jihvā - the sense of taste, experienced upon the tongue; iti - thus; jñāna-śaktayaḥ - the knowledge-acquiring senses; vāk - speech; pāṇi - the hands; upastha - the genitals; pāyu - the anus; aṅghriḥ - and the legs; karmāṇi - the working senses; aṅga - My dear Uddhava; ubhayam - belonging to both these categories; manaḥ - the mind.

Hearing, touch, sight, smell and taste are the five knowledge-acquiring senses, My dear Uddhava, and speech, the hands, the genitals, the anus and the legs constitute the five working senses. The mind belongs to both these categories.