sañcārī - passing; sāttvika - natural; sthāyi - continuously existing; bhāvera - of ecstasies; lakṣaṇa - symptoms; mukhe - in the facial expressions; netre - in the movement of the eyes; abhinaya - the dramatic performance; kare prakaṭana - he demonstrates.

He taught them how to express the symptoms of continuous, natural and transitional ecstasies with the movements of their faces, their eyes and the other parts of their bodies.