guṇa-ādhikye - by the increase of transcendental qualities; svāda-ādhikya - increase of taste; bāḍe - increases; prati-rase - in each mellow; śānta - of neutrality; dāsya - of servitude; sakhya - of fraternity; vātsalyera - and of parental affection; guṇa - the qualities; madhurete - in the conjugal mellow; vaise - appear.

“As the qualities increase, so the taste also increases in each and every mellow. Therefore the qualities found in śānta-rasa, dāsya-rasa, sakhya-rasa and vātsalya-rasa are all manifested in conjugal love [mādhurya-rasa].